quill in WordNet English dictionary

1. the hollow spine of a feather
   quill  calamus  shaft 
   
2. any of the larger wing or tail feathers of a bird
   flight feather  pinion  quill  quill feather 
   
3. a stiff hollow protective spine on a porcupine or hedgehog
   quill 
   
4. pen made from a bird's feather
   quill  quill pen